To put it in basic terms the coolant flows along block & around cyls from front of block to rear, then up thru rear of head gasket & along cyl head to front & exits via T/ Stat.
So there is a temperature gradient in that circuit..... when doing race jet boats we tried to keep that gradient within a 30° C window, any more than that it created havoc with piston to bore and other problems in relation to shut down & restart. Cars once warmed up dont [ or should not ] experience the wild temp swings that boats do in those circumstance's. The bleeds at rear that GT40's etc seem to require are just that -steam or air bleeds. I think most GT 40's built with an engine install that has the front of the engine elevated slightly could possibly get away without the rear bleeds, but they are handy if the motor is in a race situation to get rid of steam pockets etc