I do not agree on this one, and most manufacturers will advice against running a carb backwards and most carb related books do say also.
Primary bowl always up front, cause when accelerating the fuel sloshes to the back, and so on into the primary metring system with the most importance, it keeps the emulsion tubes filled. (main jet - emulsion tube - air correction jet)
Running a carb backwards, on accelerating will slosh the fuel from the primary system to the back of the float chamber... out of the carbs fuel system, lowering the fuel level of the emulsion tubes, creating a lean mixture. (flat spot, hesitation, bogging, misfire, etc).
These issues won't show up on the dyno as the car stands still.
For fast accelerating cars Holley offers jet extensions for the secondary float chamber to keep the jets fueled. But thats for the secondary float chamber only.
I made myself a long enough throttle cable ( & a spare one ) to run the carb the right way around.